Rain beetle

Family of beetles

Rain beetles
Temporal range: Late Cretaceous–recent
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Pleocoma
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Scarabaeiformia
Superfamily: Scarabaeoidea
Family: Pleocomidae
LeConte, 1861
Genera
  • Pleocoma LeConte, 1856
  • Cretocoma Nikolaev, 2002
  • Proteroscarabeus Grabau 1923

The rain beetles are a group of beetles whose extant species are found only in the far west of North America.[1] They spend most of their lives underground, emerging in response to rain or snow, thus the common name. Formerly classified in the Scarabaeidae (and later the Geotrupidae), they are currently assigned to their own family Pleocomidae, considered the sister group to all the remaining families of Scarabaeoidea. The family contains a single extant genus, Pleocoma, and two extinct genera, Cretocoma, described in 2002 from Late Cretaceous deposits in Mongolia,[2] and Proteroscarabeus of Late Cretaceous China.[3]

Rain beetles feed on various deciduous fruit trees, feeding on their roots, resulting in patches or winding bands on tree roots. This ultimately causes reduced foliage and fruit yield for trees. The larvae of rain beetles can only be found through digging to the root zone where larvae feed and observing damages. Male Pleocoma beetles, however, are attracted to light and can be monitored through that method.[4]

Male rain beetles exhibit most mating behavior during winter nights. As the females are flightless, the males spend a large portion of time walking on snow in search of them. The male's ability to maintain a high internal temperature is critical to their mating and survival prospects, as their thoracic pile is ineffective insulation.[5]

Extant members of Pleocoma are known from extreme southern Washington, throughout the mountains of Oregon and California, and into the extreme north of Baja California.[1]

References

  1. ^ a b Hovore, Frank T. "Generic Guide to New World Scarab Beetles". unsm-ento.unl.edu. University of Nebraska State Museum - Division of Entomology. Retrieved 14 December 2020.
  2. ^ Frank-Thorsten Krell. "Catalogue of fossil Scarabaeoidea (Coleoptera: Polyphaga) of the Mesozoic and Tertiary" (PDF). Natural History Museum. Archived from the original (PDF) on July 20, 2011. Retrieved March 7, 2011.
  3. ^ Krell, Frank-Thorsten. "The fossil record of Mesozoic and Tertiary Scarabaeoidea (Coleoptera: Polyphaga)." Invertebrate Systematics 14.6 (2000): 871-905.
  4. ^ "Rain Beetles | WSU Tree Fruit | Washington State University". Retrieved 2024-03-16.
  5. ^ Morgan, Kenneth R. (1987-03-01). "Temperature Regulation, Energy Metabolism and Mate-Searching in Rain Beetles (Pleocoma Spp.), Winter-Active, Endothermic Scarabs (Coleoptera)". Journal of Experimental Biology. 128 (1): 107–122. doi:10.1242/jeb.128.1.107. ISSN 0022-0949.

External links

  • iconInsects portal
  • Data related to Pleocomidae at Wikispecies
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Extant Coleoptera families
Suborder Archostemata
  • Crowsoniellidae (Crowsoniella relicta)
  • Cupedidae (reticulated beetles)
  • Jurodidae (Sikhotealinia zhiltzovae)
  • Micromalthidae
  • Ommatidae
Suborder Adephaga
Extant families
  • Amphizoidae (trout-stream beetles)
  • Aspidytidae
  • Carabidae (ground beetles)
  • Cicindelidae (tiger beetles)
  • Dytiscidae (predaceous diving beetles)
  • Gyrinidae (whirligig beetles)
  • Haliplidae (crawling water beetles)
  • Hygrobiidae
  • Meruidae (Meru phyllisae)
  • Noteridae (burrowing water beetles)
  • Trachypachidae (false ground beetles)
Suborder Myxophaga
Suborder Polyphaga
Bostrichiformia
Bostrichoidea
  • Bostrichidae (auger beetles)
  • Dermestidae (skin beetles)
  • Endecatomidae
  • Jacobsoniidae (Jacobson's beetles)
  • Nosodendridae (wounded-tree beetles)
  • Ptiniidae (furniture beetles, death watch beetles, spider beetles)
Derodontoidea
  • Derodontidae (tooth-necked fungus beetles)
Cucujiformia
Chrysomeloidea
Cleroidea
Coccinelloidea
Cucujoidea
Curculionoidea
(weevils)
  • Anthribidae (fungus weevils)
  • Attelabidae (leaf-rolling weevils)
  • Belidae (primitive weevils)
  • Brentidae (straight snout weevils, New York weevil)
  • Caridae
  • Curculionidae (true weevils, bark beetles, ambrosia beetles)
  • Nemonychidae (pine flower weevils)
Lymexyloidea
  • Lymexylidae (ship-timber beetles)
Tenebrionoidea
  • Aderidae (ant-like leaf beetles)
  • Anthicidae (ant-like flower beetles)
  • Archeocrypticidae (cryptic fungus beetles)
  • Boridae (conifer bark beetles)
  • Chalcodryidae
  • Ciidae (minute tree-fungus beetles)
  • Melandryidae (false darkling beetles)
  • Meloidae (blister beetles)
  • Mordellidae (tumbling flower beetles)
  • Mycetophagidae (hairy fungus beetles)
  • Mycteridae (palm and flower beetles)
  • Oedemeridae (false blister beetle)
  • Perimylopidae, or Promecheilidae
  • Prostomidae (jugular-horned beetles)
  • Pterogeniidae
  • Pyrochroidae (fire-coloured beetles)
  • Pythidae (dead log bark beetles)
  • Ripiphoridae (wedge-shaped beetles)
  • Salpingidae (narrow-waisted bark beetles)
  • Scraptiidae (false flower beetles)
  • Stenotrachelidae (false longhorn beetles)
  • Synchroidae (synchroa bark beetles)
  • Tenebrionidae (darkling beetles)
  • Tetratomidae (polypore fungus beetles)
  • Trictenotomidae
  • Ulodidae
  • Zopheridae (ironclad beetles, cylindrical bark beetles)
Elateriformia
Buprestoidea
  • Buprestidae (jewel beetles, or metallic wood-boring beetles)
  • Schizopodidae
Byrrhoidea
  • Byrrhidae (pill beetles)
  • Callirhipidae (cedar beetles)
  • Chelonariidae (turtle beetles)
  • Cneoglossidae
  • Dryopidae (long-toed water beetles)
  • Elmidae (riffle beetles)
  • Eulichadidae (forest stream beetles)
  • Heteroceridae (variegated mud-loving beetles)
  • Limnichidae (minute mud beetles)
  • Lutrochidae (travertine beetles)
  • Psephenidae (water-penny beetles)
  • Ptilodactylidae
Dascilloidea
  • Dascillidae (soft bodied plant beetles)
  • Rhipiceridae (cicada beetle, cicada parasite beetles)
Elateroidea
  • Artematopodidae (soft-bodied plant beetles)
  • Brachypsectridae (Texas beetles)
  • Cantharidae (soldier beetles)
  • Cerophytidae (rare click beetles)
  • Elateridae (click beetles)
  • Eucnemidae (false click beetles)
  • Jurasaidae
  • Lampyridae (fireflies)
  • Lycidae (net-winged beetles)
  • Omethidae (false fireflies, long-lipped beetles)
  • Phengodidae (glowworm beetles)
  • Rhagophthalmidae
  • Sinopyrophoridae
  • Throscidae (false metallic wood-boring beetles)
Rhinorhipoidea
  • Rhinorhipidae (Rhinorhipus tamborinensis)
Scirtoidea
Scarabaeiformia
Scarabaeoidea
  • Belohinidae (Belohina inexpectata)
  • Bolboceratidae
  • Diphyllostomatidae (false stag beetles)
  • Geotrupidae (dor beetles)
  • Glaphyridae (bumble bee scarab beetles)
  • Glaresidae (enigmatic scarab beetles)
  • Hybosoridae (scavenger scarab beetles)
  • Lucanidae (stag beetles)
  • Ochodaeidae (sand-loving scarab beetles)
  • Passalidae (betsy beetles)
  • Pleocomidae (rain beetles)
  • Scarabaeidae (scarabs)
  • Trogidae (hide beetles)
Staphyliniformia
Histeroidea
  • Histeridae (clown beetles)
  • Sphaeritidae (false clown beetles)
  • Synteliidae
Hydrophiloidea
Staphylinoidea
  • Agyrtidae (primitive carrion beetles)
  • Hydraenidae
  • Leiodidae (round fungus beetles)
  • Ptiliidae (feather-winged beetles)
  • Silphidae (carrion beetles)
  • Staphylinidae (rove beetles)
Taxon identifiers
Pleocomidae